On Stalin's orders Vladimir Gregorovich Yermolaev began design work on a long-range bomber, the DB-240 prototype of which first flew in
Yermolajev Yer-2 (DB-240)
1940. This twin diesel-engined low-wing monoplane was based on the design of R. L. Bartini's Stal' 7 and had the same distinctive inverted-gull wings. Designated Yer-2, more than 400 were built 1940- 1944 and used principally as long-range night bombers. A Yer-20N special-purpose long-range transport version, which carried 18 passengers, was developed from the bomber.

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